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1 March 2005 Oryctoantiquus borealis, New Genus and Species from the Eocene of Oregon, U.S.A., the World's Oldest Fossil Dynastine and Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)
Brett C. Ratcliffe, Dena M. Smith, Diane Erwin
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Abstract

Oryctoantiquus borealis Ratcliffe and Smith is described as a new genus and new species of fossil dynastine Scarabaeidae from West Branch Creek of the Middle Eocene Clarno Formation (44.6–46.8 Ma) of north-central Oregon, U.S.A. This specimen is the oldest specimen attributable to the subfamily Dynastinae and the largest fossil in the family Scarabaeidae.

Brett C. Ratcliffe, Dena M. Smith, and Diane Erwin "Oryctoantiquus borealis, New Genus and Species from the Eocene of Oregon, U.S.A., the World's Oldest Fossil Dynastine and Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)," The Coleopterists Bulletin 59(1), 127-135, (1 March 2005). https://doi.org/10.1649/0010-065X(2005)059[0127:OBNGAS]2.0.CO;2
Received: 16 June 2004; Accepted: 1 December 2004; Published: 1 March 2005
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