Jing Ren,¹ Natasha de Gunten,² Alexander S. Konstantinov,³ Fredric V. Vencl,⁴ Siqin Ge,^1,* David L. Hu<sup>2,5,**

1</sup>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology Chinese Academy of Sciences, Beijing 100101, China
²Woodruff School of Mechanical Engineering, Georgia Institute of Technology, Georgia 30332, USA
³Systematic Entomology Laboratory, ARS, USDA, c/o Smithsonian Institution, National Museum of Natural History, Washington DC 20013, USA
⁴Department of Ecology and Evolution, Stony Brook University, Stony Brook, New York 11794, USA
⁵School of Biology, Georgia Institute of Technology, Atlanta, Georgia 30332, USA

^*gesq@ioz.ac.cn (SG)
^**Corresponding author. E-mail: hu@me.gatech.edu (DLH)