Michio Hori,¹ Mifuyu Nakajima,² Hiroki Hata,³ Masaki Yasugi,⁴ Satoshi Takahashi,⁵ Masanori Nakae,⁶ Kosaku Yamaoka,⁷ Masanori Kohda,⁸ Jyun-ichi Kitamura,⁹ Masayoshi Maehata,¹⁰ Hirokazu Tanaka,^8,11 Norihiro Okada,^12,13 Yuichi Takeuchi<sup>14,*

1</sup>1Kyoto University, Yoshida-Honmachi, Sakyo-ku, Kyoto 606-8501, Japan
²2Department of Biology, Stanford University, Stanford, CA 94305, USA
³3Graduate School of Science and Engineering, Ehime University, Bunkyocho 2-5, Matsuyama 790-8577, Ja
⁴4National Institute for Basic Biology, Higashiyama 5-1, Myodaiji, Okazaki, Aichi 444-8787, Japan
⁵5Research Group of Environmental Sciences, Nara Woman's University, Kitauoya Nishimachi, Nara 630-85
⁶6Department of Zoology, National Museum of Nature and Science, Amakubo 4-1-1, Tsukuba, Ibaraki 305-0
⁷7Kochi University, Monobe-B, Nankoku, Kochi 783-8502, Japan
⁸8Department of Biological Science, Osaka City University, Sugimotocho, Sumiyoshi-ku, Osaka 558-8585,
⁹9Mie Prefectural Museum, Isshinden-kouzubeta 3060, Tsu, Mie 514-0061, Japan
¹⁰10Kobe Gakuin University, Arise 518, Ikawadanicho, Nishi-ku, Kobe 651-2180, Japan
¹¹11Institute of Ecology and Evolution, University of Bern, Hinterkappelen 3032, Switzerland
¹²12Department of Life Sciences, National Cheng Kung University, Tainan 701, Taiwan
¹³13Foundation for Advancement of International Science, Tsukuba 305-0821, Japan
¹⁴14Department of Anatomy, Graduate School of Medicine and Pharmaceutical Sciences, University of Toya

^*Corresponding author. E-mail: takeuchi@med.u-toyama.ac.jp