Yalu Ru,^1,2,3 Ruijing Cheng,^1,4,3 Jing Shang,^1,4 Yunpeng Zhao,^1,2,* Pan Li,^1,2 Chengxin Fu<sup>1,2

1</sup>2Key Laboratory of Conservation Biology for Endangered Wildlife of the Ministry of Education, Colleg
²3Laboratory of Systematic and Evolutionary Botany and Biodiversity, Institute of Ecology and Conserv
³5These authors contributed to this work equally.
⁴4College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, People's Republic o

^*Author for correspondence: ypzhao@zju.edu.cn