Siyu Hou,^1,2,* Zhaoxia Sun,^1,* Yaoshen Li,¹ Yijie Wang,¹ Hubin Ling,¹ Guofang Xing,^1,2 Yuanhuai Han,^1,3,** Hongying Li<sup>1,***

1</sup>2College of Agriculture, Institute of Agricultural Bioengineering, Shanxi Agricultural University, T
²3Shanxi Key Laboratory of Genetic Resources and Genetic Improvement of Minor Crops, Taigu, Shanxi 03
³4Key Laboratory of Crop Gene Resources and Germplasm Enhancement on the Loess Plateau, Ministry of A

^* These authors contributed equally to this work.
^**swgctd@163.com (Y.H.H.)
^*** Authors for correspondence: hongylswgctd@163.com (H.Y.L.),