Ya Yang,^1,2,* Michael J. Moore,³ Samuel F. Brockington,⁴ Alfonso Timoneda,⁴ Tao Feng,⁴ Hannah E. Marx,^5,6 Joseph F. Walker,¹ Stephen A. Smith<sup>1

1</sup>2Department of Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, 830 North Univer
²6Current address: Department of Microbial and Plant Biology, University of Minnesota–Twin Cities, 14
³3Department of Biology, Oberlin College, 119 Woodland Street, Oberlin, Ohio 44074-1097 USA
⁴4Department of Plant Sciences, University of Cambridge, Cambridge CB2 3EA, United Kingdom
⁵5Department of Biological Sciences and Institute for Bioinformatics and Evolutionary Studies, Univer
⁶7Current address: Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, Ari

^* Author for correspondence: yangya@umn.edu