Zhi-You Guo,^1,2,* Hong-Rui Zhang,^1,3,* Nawal Shrestha,¹ Xian-Chun Zhang<sup>1,**

1</sup>2State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of
²3College of Biological Sciences and Agriculture, Qiannan Normal University for Nationalities, Guizho
³4University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China

^*These authors contributed equally to this work.
^**Author for correspondence: zhangxc@ibcas.ac.cn