Takashi Yamamoto,^1,2,* Yoshiaki Tsuda,^3,* Gustavo Maruyama Mori,^2,4 Mariana Vargas Cruz,⁴ Yoshimi Shinmura,⁵ Alison K. S. Wee,⁶ Koji Takayama,⁷ Takeshi Asakawa,⁵ Takeru Yamakawa,⁵ Monica Suleiman,⁸ Juan Núñez-Farfán,⁹ Edward L. Webb,¹⁰ Yasuyuki Watano,⁵ Tadashi Kajita<sup>1,2,**

1</sup>2United Graduate School of Agricultural Science, Kagoshima University, 1-21-24 Korimoto, Kagoshima 9
²3Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketo
³4Sugadaira Montane Research Center, University of Tsukuba, 1278-294 Sugadairakogen, Ueda, Nagano 386
⁴5Laboratório de Análise Genética e Molecular, Centro de Biologia Molecular e Engenharia Genética, Un
⁵6Department of Biology, Graduate School of Science, Chiba University, 1-33 Yayoi, Inage, Chiba, Chib
⁶7College of Forestry, Guangxi University, Nanning, Guangxi, China
⁷8Museum of Natural and Environmental History, Shizuoka, 5762 Oya, Suruga-ku, Shizuoka-shi, Shizuoka
⁸9Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota K
⁹10Departamento de Ecología Evolutiva, Instituto de Ecología, Universidad Nacional Autónoma de México
¹⁰11Department of Biological Sciences, National University of Singapore, Singapore, Singapore

^* These authors contributed equally to this work.
^**Author for correspondence: kajita@mail.ryudai.jp