Xing-Yue Ge,^1,2,* Hao Tian,^1,2,* Wan-Jin Liao<sup>1,2,**

1</sup>2State Key Laboratory of Earth Surface Processes and Resource Ecology, College of Life Sciences, Bei
²3Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, College o

^* These authors contributed equally to this work.
^** Author for correspondence: liaowj@bnu.edu.cn