We provide the first description of the nest, eggs, young, and breeding behavior of the Great Antpitta (Grallaria excelsa) in Yacambu National Park, Venezuela. The nests (n = 3) were large, bulky, open-cup structures composed of a dense assortment of live and dead mosses, rootlets, wet leaves, small stems, detritus, and live and dead fern fronds, and were lined with a thick mesh of black rootlets and rhizomorphs. Nests were built >3.8 m above the ground in live trees where dense clusters of aroid plants, epiphytes, and lianas secured them to either a vertical fork or against the trunk itself. Both adults participated in nest building; incubating two unmarked, turquoise eggs; and feeding nestlings. Mean nest attentiveness (time spent on the nest/total video time when corrected for human disturbance) was 98.8 ± 1.8% SD, and nestling feeding rates were low (one visit by each adult/5 hr total video time).